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Once there was a maiden...

...whose domain was the ages
Her wings spanned the changing seasons
Her fangs dripped the passing seconds
The day turned to night by the twisting of her tail
"What became of the Dragon of Time?"
But no one knew to ask.

The Dragon of Time

When the Primordials built Creation, and gave pieces of themselves to become the Incarnae, Haht'hek, the Dragon of Time, volunteered to oversee the Loom of Fate, which weaves the strands of the future into the tapestry of the past. The Primordials took Haht'hek and divided him up, spreading his power among his seven subsouls, turning them into Incarnae. Mercury, Venus, Mars, Jupiter, and Saturn are well known. Alongside them were Neptune and Pluto. At the moment of their creation, they spun backwards and forwards through time, their true origin wiped from all knowledge (though Jupiter may still remember). In the times before the moment of their birth they were the Fates, and from then on they were the Maidens of Destiny.

The Sons of Neptune and the Twilight Clock of Ages

When the time came to create the Exalted, the first Five Maidens each chose 20 strands of Fate to become the Fivescore Fellowship. Gaia meanwhile was creating her Elemental Dragonblooded. Neptune, having been the Reflecting/Reproductive soul of the Dragon of Time, lent her Elemental Aspect to a host of Dragonblooded, who became the Princes of Time. Pluto chose 100 of these souls for Autochthon to forge into Celestial shards, and these became the Chosen of Hours.

Pluto's Chosen were charged with keeping the Loom running smoothly through the ridiculous amounts of reality-breaking essence use that would accompany the Primordial War. The Division of Hours operated out of the Twilight Clock of Ages, a skyscraper the color of the sky at sunset with a clock face at its apex that could be seen from anywhere in Yu Shan. The Chosen of Hours were given power over healing the Loom, smoothing out the places where essence use snarled its fibers. They were both overseers and assistants to Autochthon's Pattern Spiders, easing their workload and accounting for the actions of the Fivescore Fellowship. In fact, each Chosen of Hours was matched to one of the other Chosen, similar to how each Lunar is matched to a Solar, and each Chosen of Hours was responsible for smoothing out her mate's actions in the Loom (in theory; in practice, some Sidereals took far more work to account for than others, and Chosen of Hours often found themselves sharing responsibility).

In Creation, the Sons of Neptune had similar responsibilities, though remotely. Every Solar Circle, every Airship, every Magitech Engine of Primordial Destruction was manned by at least one Son of Neptune, whose charms were designed to mitigate the damage such titanic use of Essence was sure to cause the Loom. By their protection, hopefully the war against those who created reality would manage not to destroy the machine that maintained it.

Neptune herself approached Autochthon, wishing for a ship with which she might sail through the Loom itself, along its threads of future and past so that she might better find and correct and flaws in the flow of time. Autochthon obliged her request, though in a tragic misunderstanding he used Neptune herself as a component in its construction. The Timeship was built, and Neptune was both its powering force and its guiding intelligence. Pluto permitted none but herself to pilot it, or in cases of dire emergency six Chosen of Hours could cooperatively sail the Timeship through the Loom. With the aid of the Timeship, many events that would have been too vague for astrological prophecy and thus otherwise could have caught the Exalted off guard were found and prevented.

Medicine Man: The Final Chosen of Hours